The Antonio et al. (2019) paper on Ancient Rome was released 2 and a half months ago, so I am coming a bit late for the analysis, but I had been busy before.
Using the Dodecad K12b data provided by Jovialis, I created a table of the 11 Iron Age samples from this study.
[TABLE="width: 0"]
[TR]
[TD]ID[/TD]
[TD][/TD]
[TD="align: center"]Date[/TD]
[TD]Y-DNA[/TD]
[TD="align: center"]Gedrosia[/TD]
[TD="align: center"]Siberia[/TD]
[TD]NW African[/TD]
[TD]SE Asia[/TD]
[TD]AtlanticMed[/TD]
[TD]North
Europe[/TD]
[TD]South Asia[/TD]
[TD]East
Africa[/TD]
[TD]SW Asia[/TD]
[TD]East
Asia[/TD]
[TD]Caucasus[/TD]
[TD="align: center"]SSA[/TD]
[/TR]
[TR]
[TD]R1 (Protovillanovan)[/TD]
[TD]Martinsicuro[/TD]
[TD="align: center"]930-839 BCE[/TD]
[TD][/TD]
[TD="align: center"]5.02[/TD]
[TD="align: center"]1.78[/TD]
[TD="align: center"]1.33[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]35.08[/TD]
[TD="align: center"]26.46[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.43[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]24.50[/TD]
[TD="align: center"]0.4[/TD]
[/TR]
[TR]
[TD]R1015 (Villanovan)[/TD]
[TD]Veio Grotta Gramiccia[/TD]
[TD="align: center"]900-800 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]1.56[/TD]
[TD="align: center"]0.19[/TD]
[TD="align: center"]2.85[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]47.15[/TD]
[TD="align: center"]21.66[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.46[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]20.85[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R1016 (Latin)[/TD]
[TD]Castel di Decima[/TD]
[TD="align: center"]900-700 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]3.86[/TD]
[TD="align: center"]0.98[/TD]
[TD="align: center"]1.53[/TD]
[TD="align: center"]0.19[/TD]
[TD="align: center"]47.23[/TD]
[TD="align: center"]20.37[/TD]
[TD="align: center"]1.43[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]3.31[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]21.10[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R1021 (Latin)[/TD]
[TD]Boville Ernica[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]2.11[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.96[/TD]
[TD="align: center"]0.6[/TD]
[TD="align: center"]47.70[/TD]
[TD="align: center"]24.04[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.33[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]22.26[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R435 (Latin)[/TD]
[TD]Palestrina Colombella[/TD]
[TD="align: center"]600-200 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]4.84[/TD]
[TD="align: center"]0.64[/TD]
[TD="align: center"]0.65[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]47.12[/TD]
[TD="align: center"]28.54[/TD]
[TD="align: center"]0.15[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]4.13[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]13.40[/TD]
[TD="align: center"]0.53[/TD]
[/TR]
[TR]
[TD]R437 (Latin)[/TD]
[TD]Palestrina Selicata[/TD]
[TD="align: center"]400-200 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]6.45[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]3.03[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]33.19[/TD]
[TD="align: center"]11.94[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]11.63[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]33.74[/TD]
[TD="align: center"]0.02[/TD]
[/TR]
[TR]
[TD]R473 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]1,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.68[/TD]
[TD="align: center"]0.15[/TD]
[TD="align: center"]47.26[/TD]
[TD="align: center"]22.79[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.21[/TD]
[TD="align: center"]7.39[/TD]
[TD="align: center"]0.21[/TD]
[TD="align: center"]20.17[/TD]
[TD="align: center"]0.34[/TD]
[/TR]
[TR]
[TD]R474 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]J2b[/TD]
[TD="align: center"]7.18[/TD]
[TD="align: center"]0.17[/TD]
[TD="align: center"]2.38[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]39.08[/TD]
[TD="align: center"]25.74[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.98[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]18.84[/TD]
[TD="align: center"]0.66[/TD]
[/TR]
[TR]
[TD]R475 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]13.01[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]38.12[/TD]
[TD="align: center"]12.14[/TD]
[TD="align: center"]0.37[/TD]
[TD="align: center"]1.98[/TD]
[TD="align: center"]11.26[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]22.70[/TD]
[TD="align: center"]0.42[/TD]
[/TR]
[TR]
[TD]R850 (Greek)[/TD]
[TD]Ardea[/TD]
[TD="align: center"]800-500 BCE[/TD]
[TD]T1a[/TD]
[TD="align: center"]7.30[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]4.52[/TD]
[TD="align: center"]1,00[/TD]
[TD="align: center"]21.26[/TD]
[TD="align: center"]10.54[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.43[/TD]
[TD="align: center"]14.77[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]40.10[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R851 (Latin)[/TD]
[TD]Ardea[/TD]
[TD="align: center"]800-500 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]1.59[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.39[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]49.28[/TD]
[TD="align: center"]24.97[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.14[/TD]
[TD="align: center"]1.93[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]20.26[/TD]
[TD="align: center"]0.45[/TD]
[/TR]
[/TABLE]
R437 from Palestrina Selicata appears to be an indigenous Campanian, possibly of (partial) Greek origin, that was assimilated by Latin tribes. Or someone with mixed ancestry. R850 is clearly an Aegean Greek immigrant (very similar to modern Cretans).
Analysing admixtures
What struck me at first is the presence of Mongoloid admixture in most samples (only R437, R475 and R851 lack it). This is mostly Siberian admixture, but there are also traces of East Asian and Southeast Asian. This alludes to a sort of ancient Altaic admixture. It peaks at 1.78% in the Protovillanovan individual, who is the oldest, and would be a relatively new arrival to Italy. Does that mean that the Proto-Italics possessed some kind of Siberian admixture, which spread throughout Italy and faded as they intermingled with locals? Nowadays it's so diluted as to be virtually undetectable among modern Italians (bar a few exceptions).
The second intriguing thing is the presence of Northwest African and Southwest Asian admixture in all samples. The combination of these two admixtures, combined with elevated Caucasus admixture, first made me think of the Carthaginians. Modern and Bronze Age Lebanese have high Southwest Asian and Caucasian admixture with some Gedrosian. But they also possess some Northwest African. This is surely a component that was originally Levantine and brought to Northwest Africa by the Phoenicians/Carthaginians - not actual Northwest African genes that travelled to the Near East. But why would all Iron Age Italians possess Levantine admixture, since the Phoenician did colonise the Italian peninsula (only parts of Sicily and Sardinia)?
I had a look at modern Greeks K12b admixtures. Most Greek have only traces of NW African admixture. But those from Crete (and presumably some other Aegean islands) possess 2.6% of it. They also have higher Southwest Asian admixture (14.5%) than the Greek average (10%). So could it be mixed Greek ancestry in all these samples? The Aegean-looking sample from Ardea (R850) has the most Southwest Asian (14.8%) and Northwest African (4.5%), not far from that of modern Cretans. Other samples, the proportion of Southwest Asian to Northwest African varies a lot though. Some samples have more NW African than SW Asian.
One odd case is the R475 Etruscan from Civitavecchia, displaying as much as 13% of NW African, with also considerable SW Asian (11%) and Caucasus (22.5%). As Civitavecchia is a coastal city that had trade links with North Africa in Etruscan times, I would think that this individual has mixed Carthaginian and Etruscan ancestry. This individual has also 2.5% of East Africa + Sub-Saharan Africa, and that would be more likely if that person had actual North African roots (Northwest African native mixed with Phoenician) blended with Etruscan.
Using the Dodecad K12b data provided by Jovialis, I created a table of the 11 Iron Age samples from this study.
[TABLE="width: 0"]
[TR]
[TD]ID[/TD]
[TD][/TD]
[TD="align: center"]Date[/TD]
[TD]Y-DNA[/TD]
[TD="align: center"]Gedrosia[/TD]
[TD="align: center"]Siberia[/TD]
[TD]NW African[/TD]
[TD]SE Asia[/TD]
[TD]AtlanticMed[/TD]
[TD]North
Europe[/TD]
[TD]South Asia[/TD]
[TD]East
Africa[/TD]
[TD]SW Asia[/TD]
[TD]East
Asia[/TD]
[TD]Caucasus[/TD]
[TD="align: center"]SSA[/TD]
[/TR]
[TR]
[TD]R1 (Protovillanovan)[/TD]
[TD]Martinsicuro[/TD]
[TD="align: center"]930-839 BCE[/TD]
[TD][/TD]
[TD="align: center"]5.02[/TD]
[TD="align: center"]1.78[/TD]
[TD="align: center"]1.33[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]35.08[/TD]
[TD="align: center"]26.46[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.43[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]24.50[/TD]
[TD="align: center"]0.4[/TD]
[/TR]
[TR]
[TD]R1015 (Villanovan)[/TD]
[TD]Veio Grotta Gramiccia[/TD]
[TD="align: center"]900-800 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]1.56[/TD]
[TD="align: center"]0.19[/TD]
[TD="align: center"]2.85[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]47.15[/TD]
[TD="align: center"]21.66[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.46[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]20.85[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R1016 (Latin)[/TD]
[TD]Castel di Decima[/TD]
[TD="align: center"]900-700 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]3.86[/TD]
[TD="align: center"]0.98[/TD]
[TD="align: center"]1.53[/TD]
[TD="align: center"]0.19[/TD]
[TD="align: center"]47.23[/TD]
[TD="align: center"]20.37[/TD]
[TD="align: center"]1.43[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]3.31[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]21.10[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R1021 (Latin)[/TD]
[TD]Boville Ernica[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]2.11[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.96[/TD]
[TD="align: center"]0.6[/TD]
[TD="align: center"]47.70[/TD]
[TD="align: center"]24.04[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.33[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]22.26[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R435 (Latin)[/TD]
[TD]Palestrina Colombella[/TD]
[TD="align: center"]600-200 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]4.84[/TD]
[TD="align: center"]0.64[/TD]
[TD="align: center"]0.65[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]47.12[/TD]
[TD="align: center"]28.54[/TD]
[TD="align: center"]0.15[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]4.13[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]13.40[/TD]
[TD="align: center"]0.53[/TD]
[/TR]
[TR]
[TD]R437 (Latin)[/TD]
[TD]Palestrina Selicata[/TD]
[TD="align: center"]400-200 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]6.45[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]3.03[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]33.19[/TD]
[TD="align: center"]11.94[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]11.63[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]33.74[/TD]
[TD="align: center"]0.02[/TD]
[/TR]
[TR]
[TD]R473 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]1,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.68[/TD]
[TD="align: center"]0.15[/TD]
[TD="align: center"]47.26[/TD]
[TD="align: center"]22.79[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.21[/TD]
[TD="align: center"]7.39[/TD]
[TD="align: center"]0.21[/TD]
[TD="align: center"]20.17[/TD]
[TD="align: center"]0.34[/TD]
[/TR]
[TR]
[TD]R474 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]J2b[/TD]
[TD="align: center"]7.18[/TD]
[TD="align: center"]0.17[/TD]
[TD="align: center"]2.38[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]39.08[/TD]
[TD="align: center"]25.74[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]5.98[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]18.84[/TD]
[TD="align: center"]0.66[/TD]
[/TR]
[TR]
[TD]R475 (Etruscan)[/TD]
[TD]Civitavecchia[/TD]
[TD="align: center"]700-600 BCE[/TD]
[TD]-[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]13.01[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]38.12[/TD]
[TD="align: center"]12.14[/TD]
[TD="align: center"]0.37[/TD]
[TD="align: center"]1.98[/TD]
[TD="align: center"]11.26[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]22.70[/TD]
[TD="align: center"]0.42[/TD]
[/TR]
[TR]
[TD]R850 (Greek)[/TD]
[TD]Ardea[/TD]
[TD="align: center"]800-500 BCE[/TD]
[TD]T1a[/TD]
[TD="align: center"]7.30[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]4.52[/TD]
[TD="align: center"]1,00[/TD]
[TD="align: center"]21.26[/TD]
[TD="align: center"]10.54[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.43[/TD]
[TD="align: center"]14.77[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]40.10[/TD]
[TD="align: center"]0,00[/TD]
[/TR]
[TR]
[TD]R851 (Latin)[/TD]
[TD]Ardea[/TD]
[TD="align: center"]800-500 BCE[/TD]
[TD]R1b[/TD]
[TD="align: center"]1.59[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]1.39[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]49.28[/TD]
[TD="align: center"]24.97[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]0.14[/TD]
[TD="align: center"]1.93[/TD]
[TD="align: center"]0,00[/TD]
[TD="align: center"]20.26[/TD]
[TD="align: center"]0.45[/TD]
[/TR]
[/TABLE]
R437 from Palestrina Selicata appears to be an indigenous Campanian, possibly of (partial) Greek origin, that was assimilated by Latin tribes. Or someone with mixed ancestry. R850 is clearly an Aegean Greek immigrant (very similar to modern Cretans).
Analysing admixtures
What struck me at first is the presence of Mongoloid admixture in most samples (only R437, R475 and R851 lack it). This is mostly Siberian admixture, but there are also traces of East Asian and Southeast Asian. This alludes to a sort of ancient Altaic admixture. It peaks at 1.78% in the Protovillanovan individual, who is the oldest, and would be a relatively new arrival to Italy. Does that mean that the Proto-Italics possessed some kind of Siberian admixture, which spread throughout Italy and faded as they intermingled with locals? Nowadays it's so diluted as to be virtually undetectable among modern Italians (bar a few exceptions).
The second intriguing thing is the presence of Northwest African and Southwest Asian admixture in all samples. The combination of these two admixtures, combined with elevated Caucasus admixture, first made me think of the Carthaginians. Modern and Bronze Age Lebanese have high Southwest Asian and Caucasian admixture with some Gedrosian. But they also possess some Northwest African. This is surely a component that was originally Levantine and brought to Northwest Africa by the Phoenicians/Carthaginians - not actual Northwest African genes that travelled to the Near East. But why would all Iron Age Italians possess Levantine admixture, since the Phoenician did colonise the Italian peninsula (only parts of Sicily and Sardinia)?
I had a look at modern Greeks K12b admixtures. Most Greek have only traces of NW African admixture. But those from Crete (and presumably some other Aegean islands) possess 2.6% of it. They also have higher Southwest Asian admixture (14.5%) than the Greek average (10%). So could it be mixed Greek ancestry in all these samples? The Aegean-looking sample from Ardea (R850) has the most Southwest Asian (14.8%) and Northwest African (4.5%), not far from that of modern Cretans. Other samples, the proportion of Southwest Asian to Northwest African varies a lot though. Some samples have more NW African than SW Asian.
One odd case is the R475 Etruscan from Civitavecchia, displaying as much as 13% of NW African, with also considerable SW Asian (11%) and Caucasus (22.5%). As Civitavecchia is a coastal city that had trade links with North Africa in Etruscan times, I would think that this individual has mixed Carthaginian and Etruscan ancestry. This individual has also 2.5% of East Africa + Sub-Saharan Africa, and that would be more likely if that person had actual North African roots (Northwest African native mixed with Phoenician) blended with Etruscan.